Environments during the spread of anatomically modern humans across Northern Asia 50–10 cal kyr BP: What do we know and what would we like to know?

Northern Asia (here, the Russian Federation east of the Urals) played a key role in the spread of anatomically modern humans (AMH) across the Eurasian continent during the Upper Palaeolithic (UP). This time interval witnessed the climatically harshest and most variable part of the last glacial epoch when AMH spread to all continents, with the exception of ice-covered Antarctica, thus raising questions about how humans and environments interacted. Our review of available proxy records shows that the study region was largely dominated by productive steppe and tundra plant communities, which promoted a diverse small- to mega-scale fauna throughout the UP. While this rich fauna was an ideal food resource for AMH populations, its possible influence on the growth of woody plants through grazing is less well resolved. Another non-climatic factor that may have impacted on the spread of woody taxa are human activities (e.g. setting fires to facilitate hunting). Evidence that small populations of woody taxa were distributed in climatically favourable habitats comes from plant macroremains from sediment sequences and archaeological sites and from aDNA data. Contrary to the long-standing view of a generally colder-than-present last glacial climate, these proxy records reveal evidence that summers were warmer than today by several degrees Celsius, providing additional advantages for human activities. Another benefit for large herbivores, and thus human subsistence, were the generally low winter precipitation levels (similar to those of the modern steppe regions of Mongolia), which sustained year-round grazing grounds. These factors apparently outweighed the harsh colder-than-present winter conditions and promoted habitation of AMH in Northern Asia even during the Last Glacial Maximum (LGM) ca. 30‒18 cal kyr BP. While our understanding of qualitative climate trends, mainly based on fossil pollen records, has substantially improved, quantification of climate parameters is still a challenging task. For the last glacial interval in Northern Asia, plant macroremains, chironomids, diatoms and ostracods may provide suitable alternative proxies.